Phyllotheca

Phyllotheca; Temporal range: Permian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Kingdom:	Plantae
Clade:	Embryophytes
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Subclass:	Equisetidae
Order:	Equisetales
Family:	†Phyllothecaceae
Genus:	†Phyllotheca; Brongniart (1828)
Species
	P. australis; P. brevifolia; P. griesbachii; P. indica; P. longifolia;

Phyllotheca is an extinct genus of equisetalean, related to modern horsetails. It is known from the Permian period,^[1] as well as possibly the Carboniferous.^[2] It was named was created in 1828, when Brongniart described the type species Phyllotheca australis coming from Hawkesbury River, Australia.^[3]

Species

Phyllotheca australis: Initially described by Brongniart coming from Australia.
Phyllotheca brevifolia: Described by Roesler, Boardman and Iannuzzi. Found in Paleorrota geopark on Morro Papaléo in Mariana Pimentel, Brazil. The area is in Rio Bonito Formation dating from Sakmarian in the Permian.
Phyllotheca indica: Described by Towrow in 1955. Coming from India.
Phyllotheca longifolia: Described by Roesler and Boardman. Found in Paleorrota geopark on Morro Papaléo in Mariana Pimentel, Brazil. The area is in Rio Bonito Formation dating from Sakmarian in the Permian.^[4]

Claimed records from the Jurassic of Poland are doubtful and may represent Neocalamites instead.^[5]

References

^ Neregato, Rodrigo; Boardman, Daiana Rockenbach; Rohn, Rosemarie; Iannuzzi, Roberto (2024), "Diversity and Stratigraphic Distribution of Sphenophytes in the Permian of the Paraná Basin, Brazil", in Iannuzzi, Roberto; Rößler, Ronny; Kunzmann, Lutz (eds.), Brazilian Paleofloras, Cham: Springer Nature Switzerland, pp. 579–623, doi:10.1007/978-3-030-22526-1_16, ISBN 978-3-030-22525-4, retrieved 2026-04-11
^ Djordjevic-Milutinovic, D. (2010). An overview of Palaeozoic and Mesozoic sites with macroflora in Serbia. Bulletin of the Natural History Museum, Belgrade, 3, 27–46.
^ Uma nova espécie de Phyllotheca Brongniart
^ NOVA ESPÉCIE DE SPHENOPHYTA NO EOPERMIANO DO RIO GRANDE DO SUL
^ Jarzynka, Agata; Pacyna, Grzegorz (2015-12-01). "Fossil flora of Middle Jurassic Grojec clays (southern Poland). Raciborski's original material reinvestigated and supplemented. I. Sphenophytes". Acta Palaeobotanica. 55 (2): 149–181. doi:10.1515/acpa-2015-0013. ISSN 2082-0259.

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[1] Neregato, Rodrigo; Boardman, Daiana Rockenbach; Rohn, Rosemarie; Iannuzzi, Roberto (2024), "Diversity and Stratigraphic Distribution of Sphenophytes in the Permian of the Paraná Basin, Brazil", in Iannuzzi, Roberto; Rößler, Ronny; Kunzmann, Lutz (eds.), Brazilian Paleofloras, Cham: Springer Nature Switzerland, pp. 579–623, doi:10.1007/978-3-030-22526-1_16, ISBN 978-3-030-22525-4, retrieved 2026-04-11

[2] Djordjevic-Milutinovic, D. (2010). An overview of Palaeozoic and Mesozoic sites with macroflora in Serbia. Bulletin of the Natural History Museum, Belgrade, 3, 27–46.

[3] Uma nova espécie de Phyllotheca Brongniart

[4] NOVA ESPÉCIE DE SPHENOPHYTA NO EOPERMIANO DO RIO GRANDE DO SUL

[5] Jarzynka, Agata; Pacyna, Grzegorz (2015-12-01). "Fossil flora of Middle Jurassic Grojec clays (southern Poland). Raciborski's original material reinvestigated and supplemented. I. Sphenophytes". Acta Palaeobotanica. 55 (2): 149–181. doi:10.1515/acpa-2015-0013. ISSN 2082-0259.

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